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i've got flat feet and best piece of advice i got when running is to relax your toes. (also applies when walking)
I never noticed it before but when it was brought up I found i did indeed "grip" with the toes when active (no matter what shoe i was wearing).
relaxing it helped strengthen the correct muscles and alot of my foot pain and issues went away.
might be worth a shot to see if this helps.
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